Thursday, June 5, 2025

Recent Research on Wireless Radiation and Electromagnetic Fields

I have been circulating abstracts of newly-published scientific papers on radio frequency and other non-ionizing electromagnetic fields (EMF) monthly since 2016. The complete collection contains more than 2000 abstracts with links to these papers. Several hundred EMF scientists around the world receive these updates.

To download Volume 3 which contains abstracts of papers published since 2024 
(including the new papers listed below) click on the following link (385 page pdf):

To download Volume 2 which contains abstracts of papers published from 2021 through 2023 
click on the following link (867 page pdf):

To download Volume 1 which contains abstracts of papers published from 2016 through 2020 
click on the following link (875 page pdf):

The abstracts for recently published papers appear below.

 


A comprehensive mechanism of biological and health effects of anthropogenic extremely low frequency and wireless communication electromagnetic fields

My note: This well-referenced literature review (373 refs.) explains how electromagnetic fields (EMF) produced in wireless communications--which are polarized, coherent microwaves with highly variable intensity and modulated and pulsed at extremely low frequencies--cause "cancer, infertility, electro-hypersensitivity, and various other pathologies."

Panagopoulos DJ, Yakymenko I, De Iuliis GN, Chrousos GP. A comprehensive mechanism of biological and health effects of anthropogenic extremely low frequency and wireless communication electromagnetic fields. Front. Public Health, Volume 13 - 2025 | https://doi.org/10.3389/fpubh.2025.1585441.

Abstract

Exposure to anthropogenic electromagnetic fields (EMFs), especially those of wireless communications (WC) has increased tremendously. This is an unprecedented phenomenon throughout biological evolution because, all anthropogenic EMFs, being fully polarized, coherent, and, especially WC EMFs, highly variable, differ substantially from the natural EMFs. WC EMFs consist of Microwave (MW) carrier waves, modulated, by Extremely Low Frequency (ELF) signals, and included in on/off pulses repeated at various ELF rates. Moreover, they exhibit intense random variability, mainly in the Ultra Low Frequency (ULF) band. Thus, WC EMFs are a combination of MW and ELF/ULF EMFs. The combination of polarization/coherence and intense low-frequency (ELF/ULF) variability seems to be the key to EMF-bioactivity. Epidemiological and laboratory studies highlight a connection between ELF or WC EMF exposure and cancer, infertility, electro-hypersensitivity, and various other pathologies. Studies also find DNA damage and Oxidative Stress (OS) which explain these pathologies. While man-made EMFs cannot directly ionize molecules, they are capable of doing this indirectly in biological tissue, by triggering the biosynthesis of Reactive Oxygen Species (ROS) which can damage biomolecules, including DNA. The (over)production of ROS and the consequent OS are triggered by irregular gating of Voltage-Gated Ion Channels (VGICs) in the cell membranes as described by the Ion Forced Oscillation (IFO)-VGIC mechanism: Mobile ions within VGICs forced to oscillate by the applied ELF/ULF EMFs exert forces on the voltage sensors of the VGICs, similar to or greater than the forces that physiologically gate those channels, resulting in their irregular gating (dysfunction). Dysfunction of ion channels disrupts intracellular ionic concentrations. This triggers ROS overproduction and OS by the ROS-generating systems/enzymes in the cells, such as the electron transport chain (ETC) in the mitochondria, or the NADPH/NADH oxidases (NOXs), the Nitric Oxide synthases (NOS), etc. The IFO-VGIC mechanism and the consequent OS constitute a comprehensive mechanism that explains all known adverse biological and health effects reported to be induced by anthropogenic EMFs.

Excerpts

"The recorded human and animal carcinogenicity, the DNA/genetic damage, the OS findings, and the reproductive declines due to DNA damage in ovarian or sperm cells or embryonic death, all point toward the same direction: Man-made EMFs induce OS and DNA damage, infertility, cancer, and other related pathologies. The reason why the same effects are observed in a wide variety of animals such as mammals, birds, insects, etc., and humans, is that all biological and health effects initiate in cells and all cells are essentially identical in all animals, humans, and even plants. They have identical membranes, ions, ion channels and pumps, biomolecules such as DNA, RNA, proteins, etc., water, ROS, identical cellular organelles such as nuclei, mitochondria, ribosomes, endoplasmic reticulum, etc., and very similar metabolic processes and regulatory mechanisms. These similarities at the cellular level between all animals and humans are much more fundamental than differences in volume, mass, shape, macroscopic functions, intelligence, etc. As a result, any effect induced by EMFs in animal cells such as OS, DNA damage, etc., is expected to be induced also in the human cells, and vice-versa (7, 66).

The exposure levels in the vast majority of all the aforementioned studies were significantly below the officially accepted exposure limits for ELF and RF EMFs, which are recommended by a private organization called the International Commission on Non-Ionizing Radiation Protection (ICNIRP) to prevent discharges on humans in the case of ELF and acute heating of living tissues in the case of RF/WC EMFs. It is remarkable that this organization arbitrarily ignores the overwhelming evidence of non-thermal effects which constitute the vast majority of effects of anthropogenic EMFs, and yet, governments adopt its recommendations instead of following the Precautionary Principle which dictates the obvious, that no new technology should be applied unless those who promote it have proven its safety beyond any doubt (50, 217–226).

The International Agency for Research on Cancer (IARC) branch of the World Health Organization (WHO), has, since long time, classified both ELF and “RF” (in fact WC) EMFs as possibly carcinogenic to humans (Group 2B) (204, 205, 227). Based on additional scientific evidence after the 2011 IARC classification for “RF” EMFs, several studies have argued that “RF”/WC EMFs should be re-evaluated and classified as probably carcinogenic (Group 2A) or carcinogenic (Group 1) to humans (50, 63, 64, 66, 118–120, 171, 179, 182, 183, 218, 219). Moreover, studies have asked for the urgent application of the Precautionary Principle, stricter exposure limits, especially for WC EMFs, and a moratorium on 5G rollout (50, 215, 217, 218, 225, 228)."

"In conclusion, the IFO-VGIC mechanism that explains VGIC dysfunction, and the subsequent OS, provide a comprehensive biophysical/biochemical mechanism explaining the plethora of experimental and epidemiological findings connecting anthropogenic EMF exposures with OS, DNA/cellular damage and related pathologies such as poor health, EHS, infertility, organic/neurodegenerative diseases, cancer, etc. Even though the mechanistic details of how exactly the ionic perturbations stimulate ROS production by their sources need to be further explored, the basic scheme of the complete EMF-bioeffects mechanism is revealed already. The long existing experimental and epidemiological findings connecting exposure to man-made EMFs and DNA damage, infertility, and cancer, are now explained by the presented comprehensive mechanism. We hope this provides a better understanding of the involved science, a basis for future research, and the establishment of biologically relevant EMF exposure guidelines for effective protection of public health and the environment."

Open access paper: https://www.frontiersin.org/journals/public-health/articles/10.3389/fpubh.2025.1585441/full

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Rapid Deployment of 5G Wireless Communication and Risk Assessment on Human Health: Quid Novi?

Selmaoui B, Jamal L, Michelant L. Rapid Deployment of 5G Wireless Communication and Risk Assessment on Human Health: Quid Novi? Bioelectromagnetics. 2025 May;46(4):e70005. doi: 10.1002/bem.70005. PMID: 40276940.

No abstract.

Excerpt

"... the swift and widespread rollout of 5G has not been without controversy. Alongside the excitement over its potential, significant concerns have emerged regarding its potential impact on human health. These concerns stem from the increased exposure to electromagnetic fields (EMFs) associated with 5G technology, particularly as it operates on higher frequency bands, including millimeter waves. Consequently, given the lack of publications concerning the effects of frequencies implemented for 5G (3.5–26 GHz) for the general public, more in-depth studies need to be established due to the increased debates and inconclusive reports about the subject.

Given that 5G is a relatively new technology, short- and long-term studies are still in progress to assess its health implications comprehensively. For this purpose, the European Union Commission via their institutions has launched a call for proposals in the environmental health topic (HORIZON-HL-TH-2021-ENVHLTH-02). This program was implemented to answer to the public concern about the health effect of 5G exposure. The total amount of funding was 30 million euros from Horizon Europe 2021–2027. The results should fill the current knowledge gaps on the effects of wireless technologies on health and the environment. Four projects funded by Horizon Europe have been brought together under the CLUE-H network, involving more than 60 European research organizations across four research consortia: ETAIN, GOLIAT, NextGEM, and SEAWave. Additionally, scientists from outside Europe, including the USA, Korea, and Japan, are also collaborating on these projects...."

Conclusion

"The rapid deployment of 5G brings unprecedented opportunities for technological innovation but also necessitates thorough and ongoing risk assessment regarding its potential health impacts. While current scientific consensus generally supports the safety of 5G under existing guidelines, the evolving nature of the technology, coupled with the long-term uncertainty, underscores the importance of continued research, transparent communication, and adaptive regulatory frameworks. As 5G becomes more ubiquitous, balancing its benefits with precautionary health measures will be crucial to ensuring public trust and safety."


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5G-exposed human skin cells do not respond with altered gene expression and methylation profiles

(My note: Contrary to the title of this paper and media reports, this was not a 5G study because it tested continuous waves in the millimeter band.)

Jyoti J, Gronau I, Cakir E, Hütt M-T, Lerchl A, Meyer V. 5G-exposed human skin cells do not respond with altered gene expression and methylation profiles, PNAS Nexus, 4(5), May 2025, doi: 10.1093/pnasnexus/pgaf127.

Abstract

Due to the ever-increasing wirelessly transmitted data, the development of new transmission standards and higher frequencies in the 5G band is required. Despite basic biophysical considerations that argue against health effects, there is public concern about this technology. Because the skin penetration depth at these frequencies is only 1 mm or less, we exposed fibroblasts and keratinocytes to electromagnetic fields up to ten times the permissible limits, for 2 and 48 h in a fully blinded experimental design. Sham-exposed cells served as negative, and UV-exposed cells as positive controls. Differences in gene expression and methylation due to exposure were small and not higher than expected by chance. These data strongly support the assessment that there is no evidence for exposure-induced damage to human skin cells.

This study was funded by the Bundesamt für Strahlenschutz (FKZ 3619S82470).

Open access paper: https://academic.oup.com/pnasnexus/article/4/5/pgaf127/8124427

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Impact of mobile phone-emitted non-ionizing electromagnetic radiation on parotid gland function: A comprehensive study

Jacob RA, Jose M, Pai VR, Kalal BS. Impact of mobile phone-emitted non-ionizing electromagnetic radiation on parotid gland function: A comprehensive study. International Journal of Risk & Safety in Medicine. 2025;0(0). doi:10.1177/09246479251342488.

Abstract

Background and Objective  The rapid proliferation of mobile technology has intensified debates on the potential adverse effects of non-ionizing electromagnetic radiation (NIER) from mobile phones on various human organs and cells. This study aimed to evaluate the impact of mobile phone-emitted NIER on parotid gland function.

Methodology  This cross-sectional study included 104 student volunteers from the University campus, categorized based on their mobile phone usage and exposure duration. Saliva samples were collected from the dominant (test) and non-dominant (control) sides. Key metrics measured were salivary flow rate, pH, albumin, Ischemia-Modified Albumin (IMA), and the IMA/albumin ratio (IMAR). Data analysis was performed using one-way ANOVA and Fisher’s LSD multiple comparisons, with p ≤ 0.05 considered statistically significant.

Results  The salivary flow rate and pH were higher on both sides in all groups, correlating with increased mobile phone usage duration. Although albumin levels were lower on the dominant side, they increased with longer mobile phone use. Salivary IMA and IMAR were higher on the dominant side in subjects using mobile phones for ≤3 years compared to those using them for >3 years.

Interpretation and Conclusion  Consistent exposure to NIER and the heat generated by mobile phones adversely affects parotid gland function, as indicated by increased salivary flow rate, pH, and altered levels of albumin, IMA, and IMAR. Public health recommendations should encourage reducing long-duration conversations and using earphones to minimize NIER exposure. Further studies are needed to evaluate the long-term effects of NIER on parotid gland function.

https://journals.sagepub.com/doi/10.1177/09246479251342488

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Epidemiological criteria for causation applied to human health harms from RF-EMF exposure: Bradford Hill revisited

Frank JW. Epidemiological criteria for causation applied to human health harms from RF-EMF exposure: Bradford Hill revisited. Front. Public Health, 26 May 2025. Volume 13. doi: 10.3389/fpubh.2025.1559868.

Abstract

Purpose: This paper reviews the applicability of standard epidemiological criteria for causation, to the multidisciplinary studies of RF-EMF exposure and various adverse biological and health effects, with the aim of demonstrating that these criteria, although 60 years old, are still helpful in this context—albeit in some cases not entirely straightforward to apply.

Methods: This is a commentary, based on Bradford Hill’s criteria for assessing evidence of causation, applied to recent primary studies and systematic reviews of the RF-EMF/health-effects literature. Every effort has been made to use non-epidemiological language to reach a wide readership of biologists, physicists, and engineers now active in this field.

Results: A rapidly growing number of human observational epidemiological studies have assessed the association of diverse adverse health effects with RF-EMF exposures. However, existing systematic reviews and meta-analyses of these primary studies have substantially diverged in their conclusions. The application of Bradford Hill’s epidemiological criteria for assessing evidence of causation, originally designed for use in occupational and environmental health, casts light on some of reasons for this divergence, mostly reflecting the key weaknesses in the primary literature, which are discussed in detail. As a result of these threats to their validity—particularly the facts that (1) exposure measurement is typically subject to substantial error, and (2) insufficient time has elapsed, since modern cell phone use began in earnest, to allow tumors of longer latency to develop—most primary studies to date, and therefore many published systematic reviews of them, probably underestimate the true potential for causation, if in fact this association is causal.

Conclusion and recommendations: In view of these findings, international experts representing professional and scientific organizations in this field should convene an independent Guidelines development process to inform future epidemiological studies of associations between RF-EMF exposures and human health outcomes. Wide dissemination of such Guidelines could help researchers, journals and their reviewers in this field to execute, review and publish higher-quality studies to better inform evidence-based policy.


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Skin Fibroblasts from Individuals Self-Diagnosed as Electrosensitive Reveal Two Distinct Subsets with Delayed Nucleoshuttling of the ATM Protein in Common

Sonzogni L, Al-Choboq J, Combemale P, Massardier-Pilonchéry A, Bouchet A, May P, Doré J-F, Debouzy J-C, Bourguignon M, Dréan YL, et al. Skin Fibroblasts from Individuals Self-Diagnosed as Electrosensitive Reveal Two Distinct Subsets with Delayed Nucleoshuttling of the ATM Protein in Common. International Journal of Molecular Sciences. 2025; 26(10):4792. doi: 10.3390/ijms26104792.

Abstract

Electromagnetic hyper-sensitivity (EHS) and its causal link with radio-frequencies raise a major question of public health. In the frame of the clinical study DEMETER, 26 adult volunteers self-diagnosed as EHS-positive agreed to reply to a self-assessment questionnaire and to provide a skin biopsy sampling to establish a primary fibroblast cell line. The questionnaire and the biological data revealed, independently, 2 subsets of donors associated each with a low background, highly responsive (LBHR) and a high background, lowly responsive (HBLR) phenotype. A couple of subsets based on questionnaire data and based on the yield of spontaneous DNA double-strand breaks were found to be composed of the same donors at 64% identity. After exposure to X-rays, and application of anti-γH2AX, pATM, and MRE11 immunofluorescence, all the DEMETER fibroblasts (26/26) elicited a delayed radiation-induced ATM nucleoshuttling (RIANS). The use of RIANS biomarkers showed that the 2 phenotypes described above corresponded to DEMETER donors with a high risk of cancer (LBHR) or high risk of accelerated aging (HBLR). By exposing DEMETER cells to H2O2 followed by an antioxidative agent, we confirmed that EHS may be related to the management of DNA strand breaks. A preliminary molecular model of EHS inspired by the RIANS model was proposed.

Open access paper: https://www.mdpi.com/1422-0067/26/10/4792

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Exposure Perception and Symptom Reporting in Idiopathic Environmental Intolerance Attributed to Electromagnetic Fields Using a Co-Designed Provocation Test

Ledent M, Bordarie J, Vatovez B, Dieudonné M, Prignot N, Vanderstraeten J, Bouland C, De Clercq EM. Exposure Perception and Symptom Reporting in Idiopathic Environmental Intolerance Attributed to Electromagnetic Fields Using a Co-Designed Provocation Test. Bioelectromagnetics. 2025 Apr;46(3):e70006. doi: 10.1002/bem.70006.

Abstract

Idiopathic Environmental Intolerance Attributed to Electromagnetic Fields (IEI-EMF) is a syndrome that defines people who report symptoms that they attribute to their exposure to EMF sources, without any identified underlying medical condition to explain these symptoms. To date, provocation protocols have failed to demonstrate a consistent relationship between EMF exposure and reported symptoms, raising questions among some researchers and individuals with IEI-EMF about the relevance of these protocols for studying the syndrome. To address these criticisms, a provocation protocol was co-designed in collaboration with individuals with IEI-EMF. This study presents the results of the tests, with a focus on exposure perception and symptom reporting among IEI-EMF volunteers. A total of 47 IEI-EMF volunteers were enrolled and participated in an open-field habituation session. Of these, 27 completed the first double-blind controlled exposure session, while 26 and 16 volunteers, respectively, participated in three sessions for collective analyses and 12 sessions for individual-level analyses. At the individual level, no consistent association was found between exposure perception certainty level and exposure status, except for one volunteer whose perception was mostly consistent with exposure status. Similarly, symptom reporting did not align with exposure status, except for the same volunteer, whose symptom reporting showed a borderline significant result with exposure status. However, for half of the volunteers, symptom reporting was significantly correlated with exposure perception certainty level, supporting a nocebo hypothesis. At the collective level, no consistency was observed between exposure perception certainty level, symptom reporting, and exposure status. This study discusses the conditions necessary for future provocation protocols to enhance their relevance, acceptability, and potential utility in a possible care-oriented approach. It also considers criticisms of using exposure perception and symptom reporting as outcomes in provocation protocols, despite their central role in how individuals identify themselves as individuals with IEI-EMF.


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Electrohypersensitivity: what is belief and what is known?

de Vocht F, Martin Röösli M. Electrohypersensitivity: what is belief and what is known? Frontiers in Public Health. Vol. 13. 2025. doi:10.3389/fpubh.2025.1603692.

No abstract


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What is the effect of alarmist media and radiofrequency electromagnetic field (RF-EMF) exposure on salivary cortisol and non-specific symptoms?

Verrender A, Wallace NK, Loughran SP, Wallace C, Beange J, Croft RJ (2025). What is the effect of alarmist media and radiofrequency electromagnetic field (RF-EMF) exposure on salivary cortisol and non-specific symptoms? Applied Psychology: Health and Well-Being, 17(3), e70044. doi: 10.1111/aphw.70044.

Abstract

While there is consistent evidence that the symptoms reported by people who experience Idiopathic Environmental Intolerance attributed to Electromagnetic Fields (IEI-EMF) are closely associated with a nocebo effect, and that alarmist media reports may contribute to this nocebo effect, some methodological criticisms remain to be resolved. This study aimed to replicate previous findings and determine whether viewing an alarmist media report and being openly exposed to radiofrequency electromagnetic fields (RF-EMF) could induce a salivary cortisol response. A total of 144 participants were randomly assigned to watch either an alarmist or control video before completing an open-label provocation trial where they were either exposed or not exposed to RF-EMF. Personality factors, RF-EMF risk perception (pre- and post-video), symptoms and salivary cortisol were assessed. Consistent with previous studies, participants who were aware that they were being exposed had increased symptoms compared to participants who were aware they were not being exposed. However, the current study failed to replicate an effect of viewing an alarmist media report and being openly exposed to RF-EMF on symptoms and failed to identify an effect on salivary cortisol. This suggests that awareness and belief of exposure play a more important role in symptom perception than underlying physiological processes.

CONFLICT OF INTEREST STATEMENT

Sarah P. Loughran receives funding from the National Health and Medical Research Council of Australia (NHMRC). She is the Director of Radiation Research and Advice at the Australian Radiation Protection and Nuclear Safety Agency (ARPANSA), a member of the Scientific Expert Group at the International Commission on Non-Ionizing Radiation Protection (ICNIRP), and a member of the World Health Organisation Task Group on Radiofrequency Fields and Health Risks. Rodney J. Croft receives funding from the National Health and Medical Research Council of Australia (NHMRC) and is a member of the Scientific Expert Group at the International Commission on Non-Ionizing Radiation Protection (ICNIRP). No potential competing interests were reported by the remaining authors.

Open access paper: https://iaap-journals.onlinelibrary.wiley.com/doi/full/10.1111/aphw.70044

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Exploring the impact of environmental factors on male reproductive health through epigenetics

Zhang Y, Song JY, Sun ZG. Exploring the impact of environmental factors on male reproductive health through epigenetics. Reprod Toxicol. 2025 Mar;132:108832. doi: 10.1016/j.reprotox.2025.108832. Epub 2025 Jan 6. PMID: 39778664.

Abstract

Male infertility has become an increasingly severe global health issue, with its incidence significantly rising over the past few decades. This paper delves into the crucial role of epigenetics in male reproductive health, focusing particularly on the effects of DNA methylation, histone modifications, chromatin remodeling and non-coding RNAs regulation on spermatogenesis. Exposure to various environmental factors can cause sperm DNA damage, leading to epigenetic abnormalities. Among these factors, we have discussed heavy metals (including Zinc, Cadmium, Arsenic, Copper), phthalates, electromagnetic radiation, and temperature in detail. Notably, aberrations in DNA methylation are closely associated with various symptoms of male infertility, and histone modifications and chromatin remodeling are essential for sperm maturation and function. By synthesizing existing literature and experimental data, this narrative review investigates how environmental factors influence male reproductive health through epigenetic mechanisms, thus providing new theoretical foundations and practical guidelines for the early diagnosis and treatment of male infertility.

Excerpt

3.3 Electromagnetic radiation and sperm epigenetics 

The operating frequencies for analog mobile phones are 450–900 MHz, digital phones (Global System for Mobile Communications) range from 850 to 1900 MHz, and third-generation phones operate around 2000 MHz[144]. Electromagnetic waves (EMW) can affect sperm epigenetics through two mechanisms: 1. EMW-specific effects 2. Thermal molecular effects [145].

Experiments have found that when male rats were exposed to Wi-Fi from laptops for seven hours daily, there was a reduction in sperm count and motility within a week [146,147]. Additionally, the thermal effects of electromagnetic radiation increase free radicals around the testes, which oxidize the phospholipids in the sperm outer membrane, thus reducing membrane fluidity and impairing motility [148].

Furthermore, oxidative stress mediated by electromagnetic radiation leads to the phosphorylation of heat shock proteins, causing chromatin damage [148], thereby altering sperm epigenetics. Mice lacking Hsp90 or with highly phosphorylated Hsp proteins show cessation of spermatogonia during meiosis, leading to loss of germ cells and reduced testicular size [149]. Research indicates that Hsp90 possesses ATPase activity, stimulated by nuclear autoantigenic sperm protein (NASP) in mouse sperm nuclei [150]. A deficiency in Hsp90 or high phosphorylation of Hsp proteins can halt spermatogonia during meiosis, leading to germ cell loss and reduced testicular size [149].

In vitro animal studies by Liu et al. [151] concluded that radiofrequency radiation could have detrimental effects on human semen quality. Research has found that carrying a mobile phone or Bluetooth headset in a trouser pocket or on the waist can decrease the proportion of sperm with rapid progressive motility and increase the rate of abnormal sperm morphology [152]. Therefore, men should avoid placing mobile phones near the trousers pocket or close to the testicles to prevent potential harmful effects of radiofrequency radiation on the male reproductive system [153].


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Laptop and tablet use and their influence on total motile sperm count parameters: are laptops linked to infertility in Jamaican men? 

Sterling L, Carroll K, Harris LR. Laptop and tablet use and their influence on total motile sperm count parameters: are laptops linked to infertility in Jamaican men? Rev Int Androl. 2024 Dec;22(4):25-32. doi: 10.22514/j.androl.2024.027. 

Abstract

Background: This study aims to determine the impact of laptop and tablet use on total motile sperm count (TMSC) in men being investigated for assisted reproduction.

Methods: A cross-sectional study was conducted on 156 men attending a fertility clinic in Jamaica. Routine semen analyses were performed and parameters specific to TMSC assessed. All data analyses were performed using SPSS Version 26. Logistic regression analyses were performed to independently predict the impact of quantifiable measures of laptop and tablet use. The main outcome measures were the parameters associated with TMSC.

Results: Overall, 64% of the participants reported using laptops and 36% reported using tablets. There was a significant relationship seen with time spent on laptops and time trying to conceive (p = 0.015). Regression analyses showed that persons who used their laptops for 2 to 5 h daily were approximately 16 times (adjusted Odds Ratio (aOR) = 15.9; 95% Confidence Interval (CI), 2.5-103.3, p = 0.004) more likely to be diagnosed with low semen volume (hypospermia). Although no significant association was found between total motile sperm count (TMSC) and laptop use, a trend towards significance was observed with high laptop use (p = 0.052), suggesting potential implications for TMSC as a predictor of pregnancy outcomes.

Conclusions: Our findings highlight the need for clinicians to take into consideration wireless device usage in men undergoing fertility investigations.


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Electromagnetic fields from mobile phones: a risk for maintaining energy homeostasis?

Seewooruttun C, Mai TC, Corona A, Delanaud S, Seze R, Bach V, Desailloud R, Pelletier A. Electromagnetic fields from mobile phones: a risk for maintaining energy homeostasis? Ann Endocrinol (Paris). 2025 May 6:101782. doi: 10.1016/j.ando.2025.101782.

Abstract

In the world, there is a near ubiquitous presence of a low-intensity radiofrequency electromagnetic field (RF-EMF) radiation, due to telecommunications as mobile phones. However, their rapid expansion raises concerns about possible interaction with biological mechanisms. The RF-EMF safety guidelines recommended limits to protect against the thermal heating, the most recognized effect at high intensity levels with a known biophysical mechanism. Among all the effects studied, the impact of RF-EMF exposure on thermoregulation is one of the most important aspects of this research. This review aims to present the complex relationship between RF-EMF exposure and thermoregulation, at intensity levels below the threshold to produce thermal effects. In fact, most studies showed that RF-EMF exposure at 900 MHz seems to be elicited physiological and biological effects similar to responses inducing by cold environment in two different rodent models. In this brief review, we will describe these responses observed during or after RF-EMF exposure to give physiological and molecular mechanisms of the interaction between the organism and RF-EMF exposure at low levels and the potential implications for environmental health and safety.

Conclusion

In modern society, exposure to RF-EMF is very common and nearly impossible to avoid. Despite its major impact in driving technological advancements, it is important to address the growing public concern in regards to RF-EMF exposure. The thermal effects of high-intensity RF-EMF exposure (SAR>4 W/kg) are well documented in the literature. However, there is now emerging evidence that low-intensity RF-EMF exposure (SAR<4 W/kg) can also elicit thermoregulatory responses associated with a cold sensation. The present review highlights the behavioural thermoregulation associated with a cold sensation in rodents exposed to 900 MHz. We also provide molecular insights on RF exposure effects on different thermogenic mechanisms. The first results suggest that RF exposure primarily influences WAT browning rather than BAT thermogenesis, as reported by the increased UCP1 staining and reduced adipocyte size in WAT depots. In contrast, the cold-induced transcriptional changes on BAT thermogenesis were not observed after exposure to RF, although we detected an increased plasma levels of noradrenaline, NEFA in exposed rats. In the short term, these adaptive responses do not seem to compromise homeostasis and therefore the health of organisms, but what about the long-term consequences?

With the introduction of 3.5 GHz for 5G and early use of wireless technology by infants, further research is needed to determine these thermoregulatory effects. Investigating the peripheral tail temperature and thermal preference of 5G-exposed rats at different age could provide valuable insights into their thermoregulatory responses. Additionally, the implication of the thyroid on these responses need to be more investigate since this organ could allow a long-term adaptative response.


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Potential Impacts of Radiofrequency Electromagnetic Fields on the Central Nervous System, Brain Neurotransmitter Dynamics and Reproductive System

Eskandarpour M, Nooramin A. Potential Impacts of Radiofrequency Electromagnetic Fields on the Central Nervous System, Brain Neurotransmitter Dynamics and Reproductive System. AUT Journal of Electrical Engineering, 2025. doi: 10.22060/eej.2025.23753.5631.

Abstract

Human life has been increasingly affected by the rapid advancement of electronic technology and the widespread use of devices emitting electromagnetic radiation (EMR), such as Wi-Fi and mobile phones. While much remains unclear, studies suggest that electromagnetic fields (EMFs) can influence human health, particularly reproduction and the nervous system. EMF exposure, including from non-ionizing radiation produced by Wi-Fi and mobile phones, has been linked to potential effects on the male and female reproductive systems, embryonic development, and neuronal health. Key mechanisms include oxidative stress, thermal effects, changes in neurotransmitter metabolism, receptor function, nerve cell apoptosis, and ion channel dynamics. However, the long-term health risks, especially in children and adolescents due to prolonged exposure, remain a topic of debate. Despite current studies not confirming that RF-EMW from Wi-Fi exceeds safety guidelines, further research is essential to fully understand the implications of RF-EMW exposure on human health, particularly regarding reproduction and neurological effects. This review highlights the need for updated safety standards, more refined regulatory frameworks, and long-term investigations to clarify the potential biological and neurobiological consequences of EMF exposure.

Conclusion

RF-EMR is a form of nonionizing radiation that does not have the energy to destroy biological structures directly. However, it impressively and importantly acts on the human nervous system. Although RF-EMR itself does not act by ionizing molecules, the exposure to it can further bring about thermal effects at higher frequencies whereby raised temperature of the tissue may have an impact on cellular processes. Other than thermal effects, RF-EMR exposure can cause non-thermal biological responses through mechanisms such as oxidative stress, altered neurotransmitter activity, and changes in the flux of calcium ions within neurons. These cellular disruptions can lead to neuronal apoptosis, which hampers appropriate brain function and can potentially affect cognition and behavior. While the effects on human health are still debated, it is evident that the nervous system, given its sensitivity to electromagnetic disturbances, may be vulnerable to prolonged exposure. While radiofrequency electromagnetic radiation from Wi-Fi, for example, does involve much smaller power compared with mobile phones, and exposure is much farther from the body, there remains considerable ignorance regarding its long-term outcomes, particularly to sensitive populations of children and teenagers.

While various studies have tended to address a number of questions about RF-EMR risk, especially for brain function and reproductive health, current evidence remains conflicting and thus has necessitated further detailed investigation. It is not possible to decide on a definite conclusion because of the complexity of the biological systems implicated and the difficulty in extrapolating from animal models to human health. However, while neutral or even positive effects can be envisaged at specific levels of RF-EMR exposure, particularly in medical uses, there is still concern about possible risks that may occur above threshold values, particularly in the nervous and reproductive systems. These are potentially very disruptive to neurotransmitter functions, neuronal health, and even sperm and egg cell integrity. Larger, more comprehensive, long-term studies are urgently needed to ascertain the level of exposure that is not harmful and for a full understanding of all biological effects. In summary, exposure to RF-EMR continues to increase with each new generation of technology, including 5G. It is, therefore, of crucial importance that accurate safety guidelines and regulatory frameworks are updated. Continued research into the possible health risks, in particular the cumulative effects of long-term exposure, is essential to safeguarding public health, especially for the younger generation who are exposed more frequently and for longer periods.

Open access paper: https://eej.aut.ac.ir/article_5734.html

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Assessing RF EMF exposure in multiple microenvironments across ten European countries with a focus on 5G

Veludo AF, Stroobandt B, Van Bladel H, Sandoval-Diez N, Deprez K, Aerts S, ..., de Vocht F, Joseph W, Röösli M. Assessing radiofrequency electromagnetic field exposure in multiple microenvironments across ten European countries with a focus on 5G. Environment International, 2025. doi: 10.1016/j.envint.2025.109540.

Abstract

To evaluate the implementation of 5G in Europe, we have systematically measured environmental, auto-induced downlink (DL) and uplink (UL) radiofrequency electromagnetic field (RF-EMF) exposure in more than 800 microenvironments in ten European countries. Outdoor, indoor, and public transport microenvironments were measured in two cities and three villages in each country. Exposure was measured during three mobile-phone user scenarios: flight mode (non-user), inducing maximum DL traffic (max DL) or maximum UL traffic (max UL). The mobile phone was carried in a backpack, placed 30 cm from an ExpoM-RF 4 that continuously measured 35 frequency bands (87.5 MHz–6 GHz). For each user scenario, mean exposure levels were calculated. In the non-user scenario, mean exposure levels ranged from 0.33 to 1.72 mW/m2 per country and were lower in Switzerland, Belgium and Italy. RF-EMF levels were, on average, 80 % lower in villages compared to cities, with DL bands contributing the most in this scenario. During max DL, exposure increased mainly due to the 5G band at 3.5 GHz (mean exposure per country 2.61–11.12 mW/m2). However, the time-division nature of this band prevents distinguishing between DL and UL signals with the ExpoM-RF4. Exposure levels were the highest during max UL, particularly in the Netherlands, Italy and Belgium, with 50 % of the mean levels per country above 16 mW/m2. Exposure was, on average, 35 % higher in villages compared to cities. Environmental exposure levels were below international guideline values. Countries with precautionary limits had lower environmental exposure levels but higher auto-induced uplink exposure during data upload.

Conclusions

In this comprehensive European RF-EMF measurement study, environmental exposure levels during a non-user scenario are in line with previous research in Europe and well below international guidelines, suggesting that environmental exposure has not substantially changed since the introduction of 5G. Moreover, we could demonstrate that environmental RF-EMF exposure is mainly attributed to the Mobile DL frequency bands emitted from base stations and increases with increased urbanisation, a result of the dense network in urban settings. Additionally, we observed that environmental exposure levels are generally lower in countries with precautionary limits, namely Switzerland, Italy, and Belgium. Inducing DL traffic resulted in higher personal exposure levels, attributed to the Mobile TDD 3.5 GHz in countries where 5G is widely available. However, more studies are needed to understand the real contribution of beamforming when inducing traffic. RF-EMF exposure from an own phone during data upload was attributed mainly to 4G Mobile UL bands, and to a lower extent to the 5G band at 3.5 GHz. Additionally, in this scenario exposure levels decreased with increasing urbanisation.

Future research should monitor the evolution of environmental and auto-induced RF-EMF exposure in the different countries alongside the ongoing deployment of 5G networks in Europe.


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5G EMF Exposure at 3.6 GHz in Greece Using Data From Frequency-Selective Monitoring Sensors

Iakovidis S, Manassas A, Apostolidis C, Samaras T. 5G EMF Exposure at 3.6 GHz in Greece Using Data From Frequency-Selective Monitoring Sensors. Bioelectromagnetics. 2025 May;46(4):e70008. doi: 10.1002/bem.70008. 

Abstract

The introduction of 5G networks has raised public concerns about potential changes in environmental electromagnetic field (EMF) exposure. This study analyzes continuous monitoring data collected over 2 years (August 2022-October 2024) from 13 frequency-selective monitoring sensors located in Greece's five largest cities. Focusing on the 3.6 GHz band, we evaluated trends and weekly variations in EMF levels. Results indicated a gradual increase in EMF exposure at 3.6 GHz, driven by the growing penetration of 5G infrastructure and devices. Notably, this band exhibited higher maximum-to-median power density ratios compared to other frequency bands, attributable to active antenna systems' characteristics and traffic variations. Applying the ICNIRP 2020 guidelines, we found that 30-min averaged values significantly reduced these variations. All measured EMF levels, including maximum values, remained well below Greek and international safety limits. These findings, especially the increasing trend identified for the EMF levels, underscore the importance of continuous monitoring networks for assessing EMF exposure to existing and emerging telecommunications networks and ensuring compliance with safety standards.

Summary

  • Over the last 2 years, 5G EMF levels at the 3.6 GHz frequency band increased due to network expansion but stayed well below ICNIRP and Greek reference levels.

  • The 3.6 GHz band showed much higher maximum-to-median E-field variability than other frequency bands, driven by active antenna systems and traffic variations.

  • Using 30-min averaging, as recommended by ICNIRP (2020), significantly reduced exposure variability at 3.6 GHz, aligning it with the one observed in frequency bands used by cellular networks of previous generations.

Excerpts

"A more representative view of all 13 monitoring sensors used in this study is provided in Figure 4. The figure shows clearly that in the 3.6 GHz frequency band the ratio of the maximum 6-min-averaged power density in a week can become 12 times higher than the median 6-min-averaged power density of that week."

"The maximum-to-median ratio for the 30-min averaged values at the 3.6 GHz frequency band, became of the same order of magnitude (≈3) as those at frequency bands of older generation cellular networks (≈2)."

"The highest 6-min-averaged power density registered over 2 years was 0.053 W/m2, which is much lower than both the ICNIRP and the Greek reference levels for general public exposure, that is, 10 and 6 W/m2, respectively. It's worth mentioning that any E-field value used in this study corresponds to the 6-min RMS, that is, it is immediately comparable to established safety limits in Greece. However, the significantly higher E-field variability at the 3.6 GHz frequency band, compared to other cellular network bands, highlights the need for application of “extrapolation to the maximum” techniques when 6-min spot (in situ) measurements are performed for compliance assessment. This point should also be considered when exposure assessment studies are performed."

"Conclusions

In this study, continuous measurements of E-field from 13 monitoring sensors located in the five largest Greek cities and collected over the last 2 years were analyzed with a focus on the 3.6 GHz frequency band, where 5G networks operate. An increasing trend in environmental EMF levels at this band over these years was identified but the maximum registered incident power density remained well below the reference levels of ICNIRP and the Greek legislation. A considerably higher variation of exposure levels within a week at the 3.6 GHz band compared to other frequency bands was observed in terms of weekly maximum-to-median incident power ratio. This result highlights the necessity of extrapolation to the actual maximum when short-time measurements (e.g., of 6 min) are used as a proxy for compliance assessment. However, this high maximum-to-median ratio considerably decreases by considering 30-min averaging of the measurements, as suggested by the 2020 ICNIRP guidelines. Notably, the median E-field values at the 3.6 GHz band are lower than those of legacy networks.

No significant trends at older frequency bands were identified possibly due to limited spatial sampling, with only 13 measurement locations. Therefore, further research with broader geographical coverage and additional monitoring sensors would be valuable to evaluate these trends of exposure in the context of 5G rollout."


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Deduction of Extrapolation Factors in Realistic Scenarios for In-Situ Assessment of 5G Base Stations

Villaescusa-Tebar A, Peris JR, Garcia-Pardo C. Deduction of Extrapolation Factors in Realistic Scenarios for In-Situ Assessment of 5G Base Stations. 2025 19th European Conference on Antennas and Propagation (EuCAP), Stockholm, Sweden, 2025, pp. 1-5, doi: 10.23919/EuCAP63536.2025.10999421.

Abstract

Accurate assessment of electromagnetic field (EMF) exposure in 5G networks presents challenges due to beamforming and advanced technologies. Traditional frequency-selective methods provide actual exposure values but cannot distinguish beams or cells. Code-selective techniques differentiate cells and beams, but require complex extrapolation processes based on unavailable network parameters like beamforming patterns. This study investigates extrapolation factors like Fextbeam in realistic indoor and outdoor 5G environments, using frequency-selective and code-selective measurements. Measurements under different traffic conditions evaluated the effect of beam patterns and power. Results showed higher variability in extrapolated values due to differences in antenna radiation patterns. Outdoor environments exhibited more stable values, indicating the critical role of antenna patterns, especially indoors.


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An approach for annual analysis of EMF exposure in highly sensitive areas of kindergartens and schools

Kljajic D, Djuric N, Pasquino N, Solmonte N, Djuric S. An approach for annual analysis of EMF exposure in highly sensitive areas of kindergartens and schools. Radiat Prot Dosimetry. 2025 May 19:ncaf047. doi: 10.1093/rpd/ncaf047.

Abstract

Continuous increase of electromagnetic field (EMF) emission in the environment causes worldwide research on various short- and long-term EMF measurements and exposure assessments. Measurements are usually performed in sensitive areas, with conventional measuring approaches and worst-case exposure assessments, which are unable to provide any deeper EMF behavior analyses. Therefore, this paper brings an innovative EMF exposure analysis approach, based on time-averaging of time series data, disclosing some concealed information on long-term exposure. The averaging was done on the set of EMF time series acquired by three sensors in the EMF Regulatory Agency for Electronic Communications and Postal Services (RATEL) monitoring network. Sensors are installed at extremely sensitive locations-on buildings of two kindergartens and an elementary school. Analyses revealed patterns of daily/weekly exposure changes and their differences between working days and weekends, as well as annual decrease/increase, at each location. Comparative analysis among locations showed differences in exposure behavior regarding its peaks, slopes of the decrease/increase, and time intervals of the day with higher exposure.

https://pubmed.ncbi.nlm.nih.gov/40384509/

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Assessment of RF EMF Exposure to Car Driver from Monopole Array Antennas in V2V Communications Considering Thermal Characteristics

My note: This paper only addresses estimated RF-EMF exposure relative to inadequate ICNIRP guidelines which only protect against heating risk.

Wang S, Lu M. Assessment of RF Electromagnetic Exposure to Car Driver from Monopole Array Antennas in V2V Communications Considering Thermal Characteristics. Sensors. 2025; 25(10):3247. doi: 10.3390/s25103247.

Abstract

Vehicles are rapidly evolving into objects of intelligent interconnection. Vehicle-to-Vehicle (V2V) communications enable the interconnection between vehicles, while also leading to new electromagnetic exposure scenarios. This paper integrates a monopole array antenna into a shark-fin antenna on the car roof for V2V communications and evaluates the specific absorption rate (SAR) and temperature rise of a human body in a smart mobility communication scenario operating at 5.9 GHz. The V2V antenna is modeled and placed on a 3D vehicle model using COMSOL Multiphysics (v.6.2) to numerically estimate the SAR in the head and body regions of the human body model (adult male) inside the vehicle. Both the localized and whole-body 30 min average SAR are lower than the International Commission on Non-Ionizing Radiation Protection (ICNIRP) occupational restrictions for electromagnetic field exposure from 100 kHz to 6 GHz, being equal in the worst-case scenario to 0.981 W/kg (for the head), which is 9.81% of the ICNIRP limit (10 W/kg), and 0.008728 W/kg (for the whole-body average), which is 2.18% of the ICNIRP limit (0.4 W/kg). The 30 min average human core temperature rise is 0.055 °C, which is 5.5% of the ICNIRP limit. This indicates that, in typical automotive scenarios, the electromagnetic exposure from a monopole array antenna for V2V communications does not pose threat to the human body. This study provides knowledge related to emerging exposure scenarios in intelligent mobility communication, which is beneficial for evaluating possible health impacts and designing public health management policies.

Open access paper: https://www.mdpi.com/1424-8220/25/10/3247

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Exposure to radiofrequency electromagnetic fields and IARC carcinogen assessment: Risk of Bias preliminary literature assessment for 10 key characteristics of human carcinogens

My note: Apparently, ICNIRP supporters plan to dismiss the IARC Group 2B classification of RF-EMF carcinogenicity and the recent WHO systematic review on animal laboratory carcinogenicity studies (Mevissen et al., 2025) by claiming that all of the in vitro and in vivo studies that found evidence for carcinogenicity are poor quality.

Simkó M, Repacholi MH, Foster KR, Mattsson M-O, Croft RJ, Scarfi MR, Vijayalaxmi. Exposure to radiofrequency electromagnetic fields and IARC carcinogen assessment: Risk of Bias preliminary literature assessment for 10 key characteristics of human carcinogens. Mutation Research - Reviews in Mutation Research, Volume 796, 2025, doi: 10.1016/j.mrrev.2025.108545.

Highlights
  • We analyzed if the ten key characteristics (KCs) of human carcinogens according to IARC are influenced by RF-EMF exposure.
  • We reviewed 159 articles by extracting relevant exposure and experimental data.
  • A risk of bias (RoB) analysis was conducted using 6 criteria.
  • There is a strong negative association between study quality and the likelihood of reporting effects of RF-EMF exposures.
  • The heterogeneity and overall poor study quality suggest the need for high-quality studies.
Abstract

This is the first assessment of evidence needed to determine whether exposure to radiofrequency electromagnetic fields (RF-EMF) exposures, below the levels recommended in the ICNIRP (2020) guidelines, can influence any of the ten key characteristics (KCs) of human carcinogens developed by the International Agency for Research on Cancer (IARC). We define the 10 KCs and their relevance to carcinogenesis; review in vivo and in vitro studies relevant to the KCs; and conduct a risk of bias (RoB) analysis using 6 criteria. We did not include KC studies on genotoxicity or oxidative stress since Romeo et al. (2024) and Meyer et al. (2024) recently published relevant systematic reviews, but note their respective conclusions. From the other 8 KCs we identified 119 in vitro and 40 in vitro measurements of in vivo studies through 30 June 2023, with 38 % reporting statistically significant effects of exposure. We identified a strong association between the quality of study and outcome, with those meeting more RoB criteria less likely to report statistically significant effects. Effects were reported over the entire frequency range, exposure levels, and biological endpoints with no apparent pattern of exposure parameters resulting in effects. Only KC10 (alters cell proliferation, cell death or nutrient supply) has sufficient studies to analyse, but the other KCs had few studies and diverse endpoints. A few relatively high-quality positive studies require follow-up through additional targeted studies. The heterogeneity and overall poor study quality suggest the need for high-quality studies on these endpoints, preferably adhering to standards such as the Organization for Economic Co-operation and Development [28].

6. Conclusions

This review aimed to assess the extent and adequacy of experimental data bearing on whether exposure to RF-EMF could influence any of the 10 IARC key characteristics of human carcinogenesis. Since two PRISMA-compliant SRs on the KCs genotoxicity and oxidative stress were recently published [35], [23], this present review was limited to the 8 remaining KCs for which no systematic reviews have been conducted so far. It became apparent that the present database of in vitro and in vivo studies was so diverse and scattered in their quality that helpful outcomes of SRs on most of the KCs would be unfeasible, and any conclusions from such reviews would have a very low confidence. The two PRISMA-compliant SRs as well as previous comprehensive reviews on in vitro and in vivo genotoxicity studies (e.g. [46], and our review of studies on the 8 KCs, have all reached the same conclusion: there is clearly a need for much higher quality RF-EMF bioeffects studies on these KCs.

There are, however, a few statistically significant results in the highest quality studies (Tier 1 studies in Romeo et al. [35] and studies meeting 5 or 6 RoB criteria (present study). These deserve close examination, and replication if warranted by stronger studies, preferably done under GLP [28], or are otherwise compliant with OHAT recommendations.

As a final comment, it is not useful for systematic reviews to examine hundreds of papers on a given topic, only to find that the overall quality of the large majority of the studies is too low to permit conclusions with a high level of confidence.

Declaration of Competing Interest

The authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper.

Acknowledgments

Authors MS, MOM, and MRS acknowledge funding from the NextGEM project from the Horizon Europe Research and Innovation programme under Grant Agreement No 101057527 for part of this study. Views and opinions expressed are those of the author only and do not necessarily reflect those of the European Union or the European Health and Digital Executive Agency. Neither the European Union nor the granting authority can be held responsible for them. The other authors received no external funding for this research.


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Electromagnetic fields regulate iron metabolism: From mechanisms to applications

Wang S, Yang J, Zhen C, Wang H, Shang P. Electromagnetic fields regulate iron metabolism: From mechanisms to applications. Journal of Advanced Research. 2025, doi: org/10.1016/j.jare.2025.04.044.

Highlights

 Electromagnetic fields (EMFs) can influence the progression of various diseases by regulating iron metabolism.
 The regulation of iron metabolism by EMFs is highly parameter-specific and varies across different tissues and cells.
 The electromagnetic properties of iron-containing molecules/proteins and iron-rich tissues influence iron metabolism modulation by EMFs.
 EMFs can modulate iron metabolism through their effects on membrane structure and ion channels.
 EMFs can modulate iron metabolism by regulating the generation and activity of Reactive Oxygen Species (ROS).

Abstract

Background  Electromagnetic fields (EMFs), as a form of physical therapy, have been widely applied in biomedicine. Iron, the most abundant trace metal in living organisms, plays a critical role in various physiological processes, and imbalances in its metabolism are closely associated with the development and progression of numerous diseases. Numerous studies have demonstrated that EMF exposure induces significant changes in both systemic and cellular iron metabolism.

Aim of Review  This review aims to summarize the evidence and potential biophysical mechanisms underlying the role of EMFs in regulating iron metabolism, thereby enhancing the understanding of their biological mechanisms and expanding their potential applications in biomedical fields.

Key Scientific Concepts of Review  In this review, we have synthesized research findings and proposed the hypothesis that the biophysical mechanisms of EMFs regulate iron metabolism involve the special electromagnetic properties of iron-containing proteins and iron-enriched tissues, as well as the modulation of membrane structure and function, ion channels, and the generation and activity of Reactive Oxygen Species (ROS). Then, the review summarizes the latest advances in the effects of EMFs on iron metabolism and their safety, as well as their impact on immunoregulation, cardiovascular diseases, neurological diseases, orthopedic diseases, diabetes, liver injury, and cancer.

Conclusion and perspective

EMFs have been widely applied as a non-invasive physical alternative or adjunctive therapy in translational medicine. The regulation of iron metabolism by EMFs is a crucial mechanism underlying their biological effects and plays a significant role in their translational applications. On one hand, iron metabolism-related indicators can be used to evaluate the biological safety of EMFs. On the other hand, the effects of EMFs on iron metabolism have potential applications in the research and treatment of various diseases, including immunoregulation, cardiovascular diseases, neurological diseases, orthopedic diseases, diabetes, liver injury, and cancer. Moreover, the synergistic regulation of iron metabolism through the combination of EMFs with drugs—particularly iron-based NPs—has notably enhanced the potential of EMFs in the biomedical field. However, conflicting results in many studies regarding the effects of EMFs on iron metabolism make it challenging to systematically summarize the regulatory patterns of EMFs on iron metabolism. Despite these inconsistencies, several general characteristics of EMF-mediated regulation of iron metabolism can still be summarized. First, the effects of EMFs on iron metabolism vary depending on the properties of the EMFs (such as intensity, gradient, frequency, etc.) and exposure time [17]. Additionally, the effects of EMFs on iron metabolism appear to exhibit a biological “window”. It has been reported that the biological effects induced by EMFs of specific intensities, frequencies, and exposure durations within this “window” may differ significantly from those outside it [123]. Furthermore, the regulation of iron metabolism by EMFs also shows significant tissue- and cell-type specificity. Compared to changes in biochemical indicators, biophysical mechanisms are typically used to provide a deeper explanation of the biological effects and patterns of EMFs. In this review, we have summarized several potential biophysical mechanisms through which EMFs regulate iron metabolism, aiming to better understand the underlying reasons for the complex regulatory characteristics of EMFs on iron metabolism, and to provide research insights for utilizing EMFs in the modulation of iron metabolism in the future. Furthermore, the descriptions of EMFs parameters and exposure forms in many studies are often vague, which limits the comparability of results across different studies. Therefore, more precise descriptions of EMF parameters and exposure methods are needed in future research on the effects of EMFs on iron metabolism and related diseases. We expect that with extensive and in-depth studies on the effects and mechanisms of EMFs in iron metabolism, the scope of EMF applications in disease prevention and treatment is likely to expand.


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Active matter as the underpinning agency for extraordinary sensitivity of biological membranes to electric fields

Mathew A, Kulkarni Y. Active matter as the underpinning agency for extraordinary sensitivity of biological membranes to electric fields. Proc Natl Acad Sci U S A. 2025 Mar 25;122(12):e2427255122. doi: 10.1073/pnas.2427255122.

Abstract

Interaction of electric fields with biological cells is indispensable for many physiological processes. Thermal electrical noise in the cellular environment has long been considered as the minimum threshold for detection of electrical signals by cells. However, there is compelling experimental evidence that the minimum electric field sensed by certain cells and organisms is many orders of magnitude weaker than the thermal electrical noise limit estimated purely under equilibrium considerations. We resolve this discrepancy by proposing a nonequilibrium statistical mechanics model for active electromechanical membranes and hypothesize the role of activity in modulating the minimum electrical field that can be detected by a biological membrane. Active membranes contain proteins that use external energy sources to carry out specific functions and drive the membrane away from equilibrium. The central idea behind our model is that active mechanisms, attributed to different sources, endow the membrane with the ability to sense and respond to electric fields that are deemed undetectable based on equilibrium statistical mechanics. Our model for active membranes is capable of reproducing different experimental data available in the literature by varying the activity. Elucidating how active matter can modulate the sensitivity of cells to electric signals can open avenues for a deeper understanding of physiological and pathological processes.

Significance

What is the minimum electric field that biological cells can detect? This question has inspired significant research as certain cells and organisms are known to possess extraordinary sensitivity to weak electrical signals. However, existing models that consider this threshold to be the electrical noise arising from equilibrium thermal fluctuations yield puzzling results that are many orders of magnitude larger than experimental measurements. We resolve this paradox by proposing a nonequilibrium statistical mechanics model in which cell membranes can exploit energy-driven active mechanisms to modulate their electrical sensitivity to sense electric fields that are far weaker than the thermal electrical noise. Our study could open avenues for harnessing the sensitivity of biological cells to electric fields for diagnostics and therapeutics.

Conclusion

In summary, we investigated the role of activity in the sensitivity of biological membranes (and cells) to electric fields by way of a nonequilibrium statistical mechanics–based model. Our study reveals that activity, attributed to different sources, can endow the membranes with the ability to sense and respond to electric fields that are far weaker than the thermal noise limit estimated from purely equilibrium statistical mechanics. Our model for active membranes is capable of reproducing different experimental data available in the literature by varying the activity and thus provides a possible resolution for a long-standing discrepancy between theoretical studies and experimental measurements for the minimum electric field that can be detected by cells. As part of future study, we intend to enrich the model to incorporate electromechanical coupling through flexoelectricity and investigate its role in the interaction of active membranes with electric fields. Understanding electromechanical biological phenomena that may be impacted by increased fluctuations due to active noise in polarization is another interesting direction of study. Elucidating how cells can modulate their interaction with electric signals through activity can proffer deeper insights into physiological processes and open avenues for applications in biotechnology and medicine.


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Single exposure to near-threshold 5G millimeter wave modifies restraint stress responses in rats

Matsumoto A, Endo I, Ijima E, Hirata A, Kodera S, Ichiba M, Tokiya M, Hikage T, Masuda H. Single exposure to near-threshold 5G millimeter wave modifies restraint stress responses in rats. Environ Health Prev Med. 2025;30:33. doi: 10.1265/ehpm.24-00321.

Abstract

Background: In response to growing concerns about the health effects of quasi-millimeter waves (qMMW) used in 5th-generation wireless systems, conservative whole-body exposure thresholds based on indirect evidence have been proposed. The guidelines define a whole-body average specific absorption rate (WBA-SAR) of 4 W/kg which causes a 1 °C increase in core temperature, as the operational threshold for adverse health effects. To address the lack of direct evidence, we recently reported that a 30-minute exposure to qMMW at 4.6 W/kg resulted in a 1 °C increase in rat core temperature. Here, we further analyzed the near-threshold stress response for the first time, using biological samples from the aforementioned and additional experiments.

Methods: A total of 59 young Sprague-Dawley rats (240-322 g) were exposed to 28 GHz for 40 minutes at WBA-SARs of 0, 3.7, and 7.2 W/kg, under normal (22.5 °C, 45-55% humidity), and heat (32 °C, 70% humidity) conditions. Rats were restrained in acrylic holders for dose control. We repeatedly measured serum and urinary biomarkers of stress response, aggregated the data, and analyzed them using a single statistical mixed model to subtract the effects of sham exposure and between-subject variation.

Results: Sham exposure induced stress responses, suggesting an effect of restraint. After the subtraction of the sham exposure effect, 28 GHz appeared to induce stress responses as evidenced by elevated serum-free corticosterone 1 or 3 days after the exposure, which was more evident in animals with a change in rectal temperature exceeding 1 °C. Urinary-free catecholamines demonstrated an inhibitory property of 28 GHz frequency exposure on the stress response as evidenced by noradrenaline on the day of exposure. Heat exposure enhanced this effect, suggesting a possible role of noradrenaline in heat dissipation by promoting cutaneous blood flow, a notion supported by the correlation between noradrenaline levels and tail surface temperature, a critical organ for heat dissipation.

Conclusions: This study is the first to demonstrate that qMMW whole-body exposure can alter the stress response as indicated by corticosterone and noradrenaline at near-threshold levels. Our findings may provide insight into the biological basis of the whole-body exposure thresholds in the international guidelines.

Open access paper: https://pmc.ncbi.nlm.nih.gov/articles/PMC12062831/

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Effects of non-ionizing radiation on the thyroid gland in rats

Sarhad ZS, Ebrahimbabaei A, Tavassoli A, Shojaeifard M. Effects of non-ionizing radiation on the thyroid gland in rats. BMC Res Notes. 2025 May 24;18(1):231. doi: 10.1186/s13104-025-07297-x.

Abstract

Objectives: This study evaluated the effects of non-ionizing electromagnetic radiation on rat thyroid function and histopathology. Forty female and thirty male Sprague-Dawley rats (200-220 g, 2 months old) were exposed to 2.45 GHz Wi-Fi, mobile jammer radiation, or a sham condition. In Group A, male rats were exposed to Wi-Fi or mobile jammers for 2 h daily for two weeks. The devices were located within a one-meter radius of the animal cage, either on or off. In the Sham group, the experimental setup was like the other groups, but the irradiating devices were turned off. Group B included non-pregnant females, pregnant (exposed and control subgroups), and their offspring. Thyroid hormones in the serum were measured, and the histology was microscopically analyzed, focusing on areas of colloid and epithelium in the thyroid follicles.

Results: Compared to the control group, T4 hormone levels were significantly different in male rats exposed to mobile jammer radiation (p-value = 0.037). In group B, significant differences were found solely in the male offspring regarding T3 levels due to jammer exposure (average = 109.00 for male offspring in the experiment vs. average = 65.50 for those in the control, p-value < 0.001). Additionally, histopathological findings indicated significant differences as well. These results highlight a potential link between exposure to electromagnetic radiation and changes in thyroid endocrine and histological parameters. Our findings suggest that ongoing assessment of existing safety guidelines on non-ionizing radiation exposure is necessary, especially concerning its effects on thyroid hormone levels and follicular histology.

Open access paper: https://bmcresnotes.biomedcentral.com/articles/10.1186/s13104-025-07297-x

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An 1800 MHz Electromagnetic Field Affects Hormone Levels, Sperm Quality, and Behavior in Laboratory Rats

Pawlak K, Bojarski B, Jagusiak W, Wojnar T, Nieckarz Z, Arent Z, Ludwiczak M, Lasko M. An 1800 MHz Electromagnetic Field Affects Hormone Levels, Sperm Quality, and Behavior in Laboratory Rats (Rattus norvegicus). Applied Sciences. 2025; 15(9):5160. doi: 10.3390/app15095160.

Abstract

In addition to natural electromagnetic fields (EMFs), so-called artificial electromagnetic fields exist in the biosphere, with mobile communications being one of their main sources. This study aimed to determine the impact of EMF at a frequency of 1800 MHz on the concentrations of selected hormones, sperm motility, viability, morphology and behaviors in laboratory rats. We used 28 rats divided into two equinumerous groups: control (n = 14) and experimental (n = 14). The rats in the experimental group were exposed to EMF for 12 weeks (for 10 min, four times daily); at the same time, the control specimens were kept in standard conditions. After 12 weeks, half of each group was killed, while the other half was maintained for another 4 weeks with no EMF emission. Elevated corticosterone levels and decreased thyroid-stimulating hormone levels were observed in the experimental specimens, which persisted for 2 weeks after the cessation of EMF emission. Exposure to EMF also resulted in decreased sperm motility and viability, as well as increased rat anxiety. This study shows that exposure to EMF (1800 MHz) may affect the endocrine status of the body and the behavior and reproductive functions of animals. However, hormonal disorders appear to be reversible.

Excerpts

"The experimental group was exposed to electromagnetic waves for 10 min, four times daily: 8:00–8:10, 14:00–14:10, 18:00–18:10, and 22:00–22:10. The source of electromagnetic radiation was a generator emitting electromagnetic waves in the frequency range of the GSM cellular network (1100–2100 MHz). The average output power of the generator was 330 mW, with an instantaneous maximum power of 2 W. The load for the power stage was a Yagi GSM antenna with omnidirectional characteristics. The distance between the field generator antenna and the animals (depending on their position in the cage) was 30–40 cm."

"The electromagnetic field emitted by the generator within the rat cages was characterized by the following parameters: the average electric field intensity ranged from 2.8 to 5.7 V/m (0.1 V/m), the average magnetic field intensity ranged from 7.2 to 14.0 mA/m (±0.2 mA/m), and the average power density ranged from 17.7 to 63.6 mW/m2 (±0.5 mW/m2) at a frequency of 1800 MHz. Throughout the experiment, the EMF background was monitored, with an average value of 0.22 ± 0.07 V/m."

Open access paper: https://www.mdpi.com/2076-3417/15/9/5160

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Impairment of Oogenesis and Folliculogenesis in Neonatal Rats after Maternal Exposure to Mobile Phones

Yousefi B, Jadidi M, Nabizadeh Z, Amjad MHT, Ardekanian M. Impairment of Oogenesis and Folliculogenesis in Neonatal Rats after Maternal Exposure to Mobile Phones. Reprod Sci. 2025 May 19. doi: 10.1007/s43032-025-01880-0.

Abstract

Today's lifestyle has led to an increase in infertility. The study investigated the effects of cell phone radiation on oogenesis, folliculogenesis, and gestation stage. Also, its impact on neonates' ovarian hormones and their ovarian development during gestation was investigated. Fifty-four virgin female Wistar rats were randomly divided into three groups: the study group (conversation and stand-by mode, n = 24), the control group (stand-by mode, n = 24), and the sham (turn off, n = 6). The study and control groups were separated into 4 subgroups, including early stage (1st week), mid-stage (2nd week), late stage (3rd week), and all stages (3 weeks). The results showed that the concentration of plasma estrogen and progesterone, ovarian primordial follicle/primary oocyte, the number of primordial follicles, and nuclei diameters in the study and control subgroups decreased significantly in comparison with the sham group. The most significant reduction was observed in subgroups in which mothers were exposed to radiation for a long period during their pregnancy. Compared to the control and sham groups, the number of primordial follicle apoptosis markedly increased in the study subgroups. The pregnant rats showed significant effects in the entire stages of pregnancy, especially during the first stage (the first week) of development, which has not been reported previously. Exposure to cell phones during the different gestation stages probably decreases ovarian hormone secretion and may harm oogenesis. It also inversely increases the apoptosis of primordial follicles. Therefore, the gestational stages and duration of exposure to cell phone radiation affect the risk of ovarian harm.

Conclusion

This study examined the effects of cell phone exposure on estrogen, progesterone, and ovarian follicles in neonatal rats immediately after birth (natal day). Compared to the sham group, the results showed that cell phone radiation in both the study and control groups caused a decrease in the number and diameter of the primordial follicle, primary oocyte, its nucleus, and the amount of estrogen and progesterone hormones in neonatal rats during different and entire weeks of gestation. Also, these radiations cause an increase in apoptosis (atretic) of primordial follicles and a reduction in the diameter of the neonates’ ovaries, which depends on the duration of radiation. Meanwhile, for the first time, the impact of mobile phones during different weeks of pregnancy was examined in this study. In sum, the most effects were observed in the third week of pregnancy. The results showed that the first week of pregnancy was also affected by cell phone effects. Although this study has investigated the effects of EMF on infertility in an animal model (rat), these findings can highlight the potential risks of exposure to mobile phone radiation during pregnancy on the future fertility of the fetus in the womb. However, further research is needed to establish a direct link between EMR exposure and infertility. Non-human primate models (monkeys) are recommended to achieve more reliable results.

Open access paper: https://link.springer.com/article/10.1007/s43032-025-01880-0

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Analyzing the Impact of Occupational Exposures on Male Fertility Indicators: A Machine Learning Approach

Mohammadi H, Khoddam S, Golbabaei F, Dehghan SF. Analyzing the Impact of Occupational Exposures on Male Fertility Indicators: A Machine Learning Approach. Reprod Toxicol. 2025 May 29:108959. doi: 10.1016/j.reprotox.2025.108959. 

Highlights

• Combines occupational epidemiology with cutting-edge machine learning for risk stratification
• Provides actionable thresholds for workplace exposure limits (e.g., WBGT ≥ 28°C linked to decreased testosterone levels)
• Bridges methodological gaps in reproductive health research through model interpretability (SHAP, feature importance analysis)

Abstract

Occupational exposures are critical factors affecting workers' reproductive health. This study investigates the impact of magnetic fields, electric fields, whole-body vibration, noise levels, and heat stress on male reproductive indicators using advanced machine learning models. The aim is to identify key risk factors and provide predictive insights into workers' reproductive health over the next decade. Data were collected from 80 male workers in an automobile part manufacturing plant, capturing demographic characteristics, occupational exposures, biochemical markers, hormone levels, and sperm parameters. Five machine learning models logistic regression, bagging classifier, extreme gradient boosting, random forest, and support vector machine were trained and evaluated using 5-fold cross-validation to determine effective predictors of reproductive health outcomes. Exposure to whole-body vibration, magnetic fields, electric fields, and heat stress closely affected free testosterone levels, with SHAP importance indicating: Magnetic Field Exposure (0.339) and Wet Bulb Globe Temperature (0.138). Worker Age (0.244) was the most influential demographic factor negatively impacting Free Testosterone. The XGBoost and random forest achieved the highest AUC (0.99), outperforming other models in predictive accuracy. The Random Forest model Importance (% Increase in MSE) predicted that Electric Field Exposure (5%) and Magnetic Field Exposure (4.7%) would have the most substantial negative impact on Free Testosterone, followed by Worker Age (4.1%). This study underscores the need for targeted interventions, such as improved workplace safety protocols and regular health monitoring, to protect workers' reproductive health.

Conclusion

This study concludes that machine learning, particularly tree-based models like Random Forest and XGBoost, can effectively identify key occupational and demographic factors influencing male reproductive health. Electric and magnetic field exposures, age, work experience, and oxidative stress biomarkers emerged as the most critical predictors. Explainable AI methods revealed complex interactions among these factors. The 10-year forecast highlighted electric field exposure as the most significant long-term risk. These findings emphasize the need for targeted interventions to reduce electromagnetic and vibration exposures and to protect aging workers.


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A Mini-Review of the Potential Health Impacts of Indoor Radiation Exposure in Companion Animals

Maipas S, Vardiambasis IO, Nikolopoulos CD, Kavantzas N. A Mini-Review of the Potential Health Impacts of Indoor Radiation Exposure in Companion Animals. Environ Health Insights. 2025 May 28;19:11786302251342972. doi: 10.1177/11786302251342972.

Abstract

Poor indoor environmental quality is a major concern for both humans and companion animals-yet its impact on the latter is often overlooked. This mini review sheds light on indoor radiation sources that may affect the health of companion animals. Radon, primarily from the ground and possibly also from cat litter, naturally occurring radionuclides in feed, radiofrequency radiation from mobile and cordless phones, Wi-Fi networks, pet tracking devices, solar radiation, and various sources of extremely low frequency radiation are all common examples. Indoor radiation, an odorless and tasteless pollutant, may have the potential to negatively impact the health and well-being of companion animals, which are involuntarily exposed to this environmental health risk factor. Therefore, preventive and precautionary measures are necessary.


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Melatonin ameliorates RF-EMR-induced reproductive damage by inhibiting ferroptosis through Nrf2 pathway activation

Wang J, Dong J, Xu Q, Yan S, Wang H, Lei H, Ma X, Yang T, Wang K, Li Z, Wang X. Melatonin ameliorates RF-EMR-induced reproductive damage by inhibiting ferroptosis through Nrf2 pathway activation. Pathol Res Pract. 2025 May 7;270:156003. doi: 10.1016/j.prp.2025.156003.

Abstract

In recent years, there has been increased attention to the deleterious impacts of radiofrequency electromagnetic radiation (RF-EMR) on male reproductive ability, necessitating the exploration of effective protective measures. Melatonin has antioxidant and anti-apoptotic effects, and there is growing evidence of its benefit to the reproductive process. However, the biochemical mechanisms by which melatonin protects against reproductive damage from RF-EMR exposure are unknown. Here, we found that prolonged (8 weeks) exposure to RF-EMR [2.45 GHz; power density, 2.5 W/m2; whole-body specific absorption rate (SAR), 0.125-0.5 W/kg] induced ferroptosis and oxidative stress in testicular tissue, leading to a decrease of sperm quality in male mice. Notably, the administration of melatonin mitigated the oxidative harm to the testicles and ferroptosis caused by RF-EMR in mice. Mechanistically, melatonin could inhibit ROS production and ferroptosis by stimulating the nuclear factor-erythroid 2-related factor 2 (Nrf2) signaling pathway through its receptors (MT1/MT2). Taken together, these results indicate that melatonin could potentially improve RF-EMR-induced reproductive damage in male mice by blocking ferroptosis through activation of the Nrf2 pathway.

Conclusions

In conclusion, this study demonstrated that RF-EMR harms male reproduction by triggering ferroptosis, a cell death process, in mouse testes. Melatonin, by activating the Nrf2 pathway via MT1/MT2 receptors, suppresses ferroptosis and protects against RF-EMR damage (Fig. 6). These findings suggest melatonin's potential clinical value. Future controlled trials will explore its efficacy in treating male infertility caused by RF-EMR exposure, making melatonin a promising candidate therapy. ...


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Exposure at 3.6 GHz in Greece Using Data From Frequency-Selective Monitoring Sensors

Iakovidis S, Manassas A, Apostolidis C, Samaras T. 5G EMF Exposure at 3.6 GHz in Greece Using Data From Frequency-Selective Monitoring Sensors. Bioelectromagnetics. 2025 May;46(4):e70008. doi: 10.1002/bem.70008.

Abstract

The introduction of 5G networks has raised public concerns about potential changes in environmental electromagnetic field (EMF) exposure. This study analyzes continuous monitoring data collected over 2 years (August 2022-October 2024) from 13 frequency-selective monitoring sensors located in Greece's five largest cities. Focusing on the 3.6 GHz band, we evaluated trends and weekly variations in EMF levels. Results indicated a gradual increase in EMF exposure at 3.6 GHz, driven by the growing penetration of 5G infrastructure and devices. Notably, this band exhibited higher maximum-to-median power density ratios compared to other frequency bands, attributable to active antenna systems' characteristics and traffic variations. Applying the ICNIRP 2020 guidelines, we found that 30-min averaged values significantly reduced these variations. All measured EMF levels, including maximum values, remained well below Greek and international safety limits. These findings, especially the increasing trend identified for the EMF levels, underscore the importance of continuous monitoring networks for assessing EMF exposure to existing and emerging telecommunications networks and ensuring compliance with safety standards.


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Terahertz Radiation Affects the Dynamics of Neurons by Decreasing Membrane Area Ratio

Ma S, Li S, Wang H, Li Y, Lu C, Li X. Terahertz Radiation Affects the Dynamics of Neurons by Decreasing Membrane Area Ratio. Brain Res Bull. 2025 May 6:111373. doi: 10.1016/j.brainresbull.2025.111373.

Abstract

Terahertz radiation at specific frequencies and energies can mediate cellular morphology or function changes by exciting nonlinear resonance effects in proteins or DNA. However, the effects of terahertz radiation on neuronal morphology and function are currently unknown, and the correlation between neuronal morphology and kinetic properties after terahertz radiation remains to be elucidated. In this paper, we first characterized the changes in neuronal morphology by the relative ratio of neuronal cytosol to protruding membrane area. Analyzed the pattern of the influence of terahertz radiation on neuronal morphology and the cumulative effect. On this basis, this paper constructs a kinetic model of neurons regulated by terahertz radiation, investigates the influence law of terahertz radiation on the kinetic properties of neurons, and analyzes the correlation between neuronal morphology and kinetic properties. The results showed that terahertz radiation caused a decrease in the membrane area ratio of neuronal cytosol to protrusion, and this effect started on the first day of terahertz radiation and lasted until the end of terahertz radiation; terahertz radiation changed the neuronal discharge pattern by decreasing the membrane area ratio of neuronal cytosol to protrusion and lowered the frequency of neuronal inter-cluster discharges and amplitude of action potentials, and increased the neuronal intra-cluster discharge. In addition, terahertz radiation can increase the peak value of neuronal postsynaptic currents by decreasing the membrane area ratio. In summary, terahertz radiation can modulate neurons' morphology and change their firing patterns and kinetic properties by affecting their morphology. These predict that terahertz radiation at specific frequencies and energies can be developed as a novel, molecular-level neuromodulation technique for intervening or treating neuronal degenerative diseases.


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Exploring the influence of Schumann resonance and electromagnetic fields on bioelectricity and human health

Nelson I. Exploring the influence of Schumann resonance and electromagnetic fields on bioelectricity and human health. Electromagn Biol Med. 2025 May 20:1-11. doi: 10.1080/15368378.2025.2508466.

Abstract

This article explores the relationship between electromagnetic fields (EMF) and biological systems, focusing on the influence of extremely low-frequency electromagnetic frequencies (ELF), particularly Schumann's resonance (SR) at 7.83 hz. Cells and proteins may have evolved to take advantage of frequencies naturally present in the Earth's EMF, potentially enhancing cellular energy levels and affecting resting membrane potential (RMP). Thus, changes in or absence of SR may have adverse effects on the functioning of the whole organism. Bioelectricity, independent of genes, has been shown to modulate health, suggesting the potential for using controlled application of EMF frequencies in treating certain types of cancer or conditions affecting the RMP. Research indicates that human brainwave activity is highly dependent on the SR, implying a correlation between atmospheric electromagnetic frequencies and brain activity. ELF, including SR, appears to modulate cellular calcium influx/efflux, likely via indirect mechanisms involving field-sensitive molecules or radical pairs that affect ion channel behavior which plays a critical role in cell signaling and regulation of various processes. It can also trigger a cascade of molecular events that ultimately lead to the generation of action potentials, affecting consciousness and behavior. The influence of atmospheric electromagnetic frequencies on human brainwave activity, modulation of cellular calcium influx/efflux, and potential effects on cellular energy levels and RMP highlight the significance of ELF in biological systems. However, further research is required to fully understand these mechanisms and their implications for human health and well-being.

Plain Language Summary

This study looks at how very low-frequency electromagnetic fields – especially the Schumann Resonance at 7.83 hz – may affect our health. These natural frequencies come from the Earth’s atmosphere and may play an important role in how our bodies and cells work. The research explores how these signals could influence the electrical activity inside our cells, including how ions like calcium move in and out. This process is essential for brain function, heart rhythm, sleep patterns, and overall health. The study suggests that human cells may have evolved to respond to these natural frequencies. Changes in, or the absence of, these signals – such as when we’re exposed to artificial electromagnetic fields from modern technology – might disrupt important processes in the body. For example, the paper highlights evidence that some low-frequency signals may help stop cancer cells from growing or improve healing, while unnatural frequencies might do harm. A key point is that our brain activity seems to align with Earth’s natural electromagnetic rhythms. This could explain why changes in these fields can affect our mood, sleep, or concentration. The study also points out that future space missions should consider simulating these natural frequencies to help astronauts stay healthy outside Earth’s environment. In short, this research helps us understand how our bodies interact with the invisible electromagnetic environment around us – and how both natural and man-made signals could influence our health in powerful ways.

Conclusions

Solar and geomagnetic activities appear to affect biological organisms in diverse ways. SR appears to be a biophysical mechanism that coordinates such a relationship.

Human EEG is highly dependent on SR, which suggests that the natural electromagnetic frequencies in the Earth’s atmosphere, known as SR, influence human brainwave activity. This suggests a correlation between atmospheric electromagnetic frequencies and brain activity.

While earlier models focused on calcium influx as the primary target of ELF fields, more recent findings indicate that weak magnetic fields likely act first on radical pairs within mitochondria. This radical pair mechanism (RPM) may influence reactive oxygen species (ROS) dynamics, which are tightly linked to intracellular calcium signaling via mitochondria-associated membranes (MAMs). These pathways can alter membrane potential, gene expression, and cellular communication.

Given this context, VGCC activation and calcium fluxes may be secondary consequences of mitochondrial redox modulation. Since calcium ions are involved in nearly all critical cellular processes – from apoptosis to signal transduction – these downstream effects can profoundly influence bioelectric regulation and health outcomes.Biological systems may have evolved to be influenced by electromagnetic frequencies in the ELF range that are naturally present in the environment. These frequencies, particularly the SR at 7.83 hz, could potentially enhance cellular energy levels and affect the RMP. Healthy cells typically have a higher RMP than those with dysfunction or disease.

During the silent phase, thalamocortical neurons might be entrained with SR, which suggests that during this phase, the electromagnetic frequencies of SR could potentially synchronize or entrain the activity of thalamocortical neurons, thereby justifying the coordination between SR and EEG.

There are ongoing studies on the potential health effects of exposure to electromagnetic fields. With the information collected from these studies, it seems plausible that unnatural EMF might affect organisms and cells in multiple ways, such as abnormal ion channels that function to coordinate the whole organism, and improper circadian rhythms.

https://www.tandfonline.com/doi/10.1080/15368378.2025.2508466

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Effect of Occupational Exposure to Low-frequency Electromagnetic Fields on Cataract Development

Validad MH, Mahjoob M, Pishjo M, Diani M, Rakhshandadi T. Effect of Occupational Exposure to Low-frequency Electromagnetic Fields on Cataract Development. J Ophthalmic Vis Res. 2025 Apr 25;20:1-6. doi: 10.18502/jovr.v20.12281.

Abstract

Purpose: Cataracts are the second leading cause of visual impairment worldwide. This study aimed to examine the impact of occupational exposure to low-frequency electromagnetic fields on cataract development.

Methods: One hundred employees of Zahedan Electricity Company participated in this study. They were assigned to four groups based on their level of exposure: regular, operational, operator personnel, and non-exposure. Based on LOCS III grading, the risk of developing different types of cataracts (i.e., nuclear, posterior subcapsular, and cortical) was evaluated for all participants.

Results: The frequency of cataracts was 62.2% in the exposure group (which includes three subgroups: the regular, operational, and operator personnel) and 53.8% in the non-exposure group. There was a significant difference between the study groups in terms of nuclear opacity grading (P = 0.003). The correlation between nuclear and posterior subcapsular cataract grading and work experience duration in the exposure group was statistically significant (P < 0.018).

Conclusion: This study's findings indicate that exposure to low-frequency electromagnetic fields such as power lines, power plants, and power distribution posts may be a risk factor for cataract development, particularly nuclear cataracts.


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Carcinogenicity of extremely low-frequency magnetic fields: A systematic review of animal studies

Luukkonen J, Roivainen P, Nieminen V, Naarala J, Mustafa E, Juutilainen J. Carcinogenicity of extremely low-frequency magnetic fields: A systematic review of animal studies. Environ Res. 2025 May 13:121819. doi: 10.1016/j.envres.2025.121819.

Abstract

Possible carcinogenicity of extremely low frequency (ELF) magnetic fields (MFs), associated with the use and transmission of electricity, has been under scientific and public debate for decades. This review aims to provide an update on studies testing carcinogenicity of ELF MFs in experimental animals. Emphasis was placed on identifying possible connections between study characteristics and the results obtained. This review followed the PRISMA (Preferred Reporting Items for Systematic Reviews and Meta-Analyses) statement, and the methodological quality of the studies was evaluated using the Risk of Bias Rating Tool for Human and Animal Studies developed by the National Toxicology Program’s Office of Health Assessment and Translation (OHAT). Publication bias was assessed using the caliper test. Fifty-four eligible studies were identified. Despite poor ratings in certain aspects of the risk of bias evaluation, the quality of the studies was generally relatively good, with only four studies receiving the weakest rating. Overall, there was very little evidence that ELF MFs alone are carcinogenic. Evidence of co-carcinogenicity, from studies that have used ELF MFs in combination with other agents, remains inconclusive. A clear indication of publication bias was observed, though it is unlikely that this bias alone explains all reported MF-modified effects. Based on the current literature, future studies on co-carcinogenicity of ELF MFs should employ approaches other than classical initiation-promotion models. Additionally, well-designed studies to better understand the reported effects of ELF MFs and the underlying mechanisms are warmly welcomed.


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Weak anthropogenic electric fields affect honeybee foraging

Mallinson VJ. Woodburn FA, O’Reilly KJ. Weak anthropogenic electric fields affect honeybee foraging. Cell Press. 2025. https://doi.org/10.1016/j.isci.2025.112550.

Highlights

• Weak anthropogenic-like electric fields reduce honeybee floral landing
• AC and positive DC fields deter honeybee landing, but negative DC does not
• Electric fields near power lines match experimental levels, spanning tens of meters
• These findings highlight electric pollution as a potential disruptor of pollination

Summary

Aerial electroreception, the detection of airborne electric fields (E-fields), is an emerging sensory system in arthropods, including bees, which can use floral E-fields as foraging cues. However, the influence of anthropogenic E-fields on these interactions remains underexplored. Through field experiments in urban meadows, we demonstrate that weak anthropogenic E-fields, including alternating current (AC) and direct current (DC) fields, significantly alter honeybee floral landing behaviors. AC and positive DC fields reduced landings by 71% and 53%, respectively, whereas negative DC fields had no statistically significant impact. Measurements of E-fields near high-voltage transmission lines revealed persistent field strengths comparable to those used experimentally, spanning tens of meters at heights relevant for bee foraging. These findings underscore the potential for anthropogenic E-fields to affect and potentially disrupt plant-pollinator interactions, threatening pollination efficiency, a cornerstone of agriculture and biodiversity. Our study highlights the need for advancing research on the ecological impacts of electric pollution.


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Effect of short-term extremely low-frequency electromagnetic field on respiratory functions 

Sirinyildiz F, Cesur G, Elmas O, Elmas S, Comlekci S, Yazıcı O, Keskin A, Keskin HE. Effect of short-term extremely low-frequency electromagnetic field on respiratory functions. Rev Assoc Med Bras (1992). 2025 Jun 2;71(4):e20241812. doi: 10.1590/1806-9282.20241812. PMID: 40465999.

Abstract

Objective: Exposure to extremely low-frequency electromagnetic fields can cause harmful or beneficial effects on living organisms. The aim of this study was to examine the potential effects of extremely low-frequency electromagnetic fields on respiratory physiology by investigating possible changes in respiratory function parameters during and after short-term extremely low-frequency electromagnetic field exposure.

Methods: Twenty Wistar albino rats were included in the study, and these rats were randomly divided into two groups: control and electromagnetic field. A noninvasive head-out plethysmography technique was used to accurately assess lung function in rats. Rats in the electromagnetic field group were exposed to electromagnetic field at a frequency of 50 Hz and a magnetic flux density of 0.3 mT for 2 min. Respiratory function parameters of both groups were recorded in three separate periods before, during and after electromagnetic field exposure. Respiratory rate, respiratory cycle duration, inspiration time, expiration time, tidal volume, minute volume, peak inspiratory flow, and peak expiratory flow were measured in these periods.

Results: There was no significant difference in the parameters measured before electromagnetic field exposure between the groups. During the electromagnetic field exposure period, the mean respiratory rate measured in the electromagnetic field group was lower compared to the control group data, while the mean respiratory cycle duration, inspiration time, and tidal volume measured in the electromagnetic field group were higher compared to the control group data. There was no significant difference in the parameters measured after electromagnetic field exposure between the groups.

Conclusion: Short-term extremely low-frequency electromagnetic field exposure decreases respiratory rate and increases respiratory cycle duration, inspiration time, and tidal volume.

Open access paper: https://www.scielo.br/j/ramb/a/9mQPKdGLX8DJ3mPwSqygwTG/?lang=en

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Electromagnetic wireless remote control of mammalian transgene expression

Lin Z, Guha Ray P, Huang J. et al. Electromagnetic wireless remote control of mammalian transgene expression. Nat. Nanotechnol. (2025). https://doi.org/10.1038/s41565-025-01929-w.

Abstract

Communication between wireless field receivers and biological sensors remains a key constraint in the development of wireless electronic devices for minimally invasive medical monitoring and biomedical applications involving gene and cell therapies. Here we describe a nanoparticle–cell interface that enables electromagnetic programming of wireless expression regulation (EMPOWER) of transgenes via the generation of cellular reactive oxygen species (ROS) at a biosafe level. Multiferroic nanoparticles coated with chitosan to improve biocompatibility generate ROS in the cytoplasm of cells in response to a low-frequency (1-kHz) magnetic field. Overexpressed ROS-responsive KEAP1/NRF2 biosensors detect the generated ROS which is rewired to synthetic ROS-responsive promoters to drive transgene expression. In a proof-of-concept study, subcutaneously implanted alginate-microencapsulated cells stably expressing an EMPOWER-controlled insulin expression system normalized blood-glucose levels in a mouse model of type 1 diabetes in response to a weak magnetic field.


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Effects of radiofrequency electromagnetic field exposure on cancer in laboratory animal studies, a systematic review

Mevissen M, Ducray A, Ward JM, Kopp-Schneider A, McNamee JP, Wood AW, Rivero TM, Straif K. Effects of radiofrequency electromagnetic field exposure on cancer in laboratory animal studies, a systematic review. Environment International. 2025, doi: 10.1016/j.envint.2025.109482.

Abstract

Background  

More than ten years ago, the World Health Organization’s (WHO) International Agency for Research on Cancer (IARC) published a monograph concluding there was limited evidence in experimental animals for carcinogenicity of Radio Frequency Electromagnetic Field (RF EMF).

Objective  

The objective of this review was to systematically evaluate the effects of RF EMF exposure on cancer in experimental animals.

Methods   

  Eligibility criteria: Based on pre-established Populations, Exposures, Comparators, Outcomes, and Study Type (PECOS) criteria, studies in experimental animals of the following study types were included: chronic cancer bioassays, initiation-(co–)promotion studies, and studies with tumor-prone animals. 
  Information sources: MEDLINE (PubMed), Science Citation Index Expanded and Emerging Sources Citation Index (Web of Science), and the EMF Portal. 
  Data abstraction and synthesis: Data are publicly available online as interactive visuals with downloadable metadata. We adapted the risk-of-bias (RoB) tool developed by Office of Health Assessment and Translation (OHAT) to include considerations pertinent to the evaluation of RF EMF exposure and cancer bioassays. Study sensitivity was assessed with a tool adopted from the Report on Carcinogens (RoC). We synthesized studies using a narrative approach. Effect size was calculated as the 1% Bayesian Average benchmark dose (BMD) of a respective study when dose–response or a trend was identified (see BMDAnalysisSupplementaryMaterial) (Supplement 1).
  Evidence Assessment: Certainty of the evidence (CoE) was assessed using the Grading of Recommendations, Assessment, Developing and Evaluations (GRADE) approach, as refined by OHAT. Evidence from chronic cancer bioassays was considered the most directly applicable to evaluation of carcinogenicity.

Results 

We included 52 studies with 20 chronic bioassays No studies were excluded based on risk of bias concerns. Studies were not considered suitable for meta-analysis due to heterogeneity in study design, species, strain, sex, exposure characteristics, and cancer outcome. No or minimal evidence of RF EMF exposure-related cancer outcomes was found in most systems or organs in any study (these included gastrointestinal/digestive, kidney, mammary gland, urinary, endocrine, musculoskeletal, reproductive, and auditory). For lymphoma (18 studies), with 6 chronic bioassays (1,120 mice, 1,780 rats) inconsistency between two chronic bioassays was not plausibly explainable, and the CoE for lymphoma was rated ‘moderate’. For brain tumors (20 studies), including 5 chronic bioassays (1,902 mice, 6,011 rats), an increase in glial cell-derived neoplasms was reported in two chronic bioassays in male rats. The CoE for an increased risk in glioma was judged as high. The BMD analysis was statistically significant for only one study and the BMD was 4.25 (95% CI 2.70, 10.24). For neoplasms of the heart (4 chronic bioassays with 6 experiments), 3 studies were performed in rats (∼2,165 animals), and 1 in mice (∼720 animals). Based on 2 bioassays, statistically significant increases in malignant schwannomas was judged as high CoE for an increase in heart schwannomas in male rats. The BMDs from the two positive studies were 1.92 (95 %CI 0.71, 4.15) and 0.177 (95 %CI 0.125, 0.241), respectively. Twelve studies reported neoplasms in the adrenal gland (5 chronic bioassays). The CoE for an increased risk in pheochromocytoma was judged as moderate. None of these findings were dose-dependent when compared to the sham controls. Sixteen studies investigated tumors of the liver with 5 of these being chronic bioassays. The CoE was evaluated as moderate for hepatoblastomas. For neoplasms of the lung (3 chronic bioassays), 8 studies were conducted in rats (∼1,296 animals) and 23 studies in mice (∼2,800 animals). In one chronic bioassay, a statistically significant positive trend was reported for bronchoalveolar adenoma or carcinoma (combined), which was rated as moderate CoE for an increase in lung neoplasms with some evidence from 2 initiation-(co–)promotion studies.

Discussion  

Meta-analysis was considered inappropriate due to the heterogeneity in study methods. The GRADE/OHAT CoE framework has not been frequently applied to animal studies and experience to date suggests refinements are needed. We deferred to standard methods in environmental health where CoE is framed in the context of strength of the evidence providing positive support for carcinogenicity. High CoE can be interpreted as the true effect is highly likely to be reflected in the apparent relationship. Moderate CoE indicates the true effect may be reflected in the apparent relationship. Cancer bioassays conducted in experimental animals are commonly used to identify potential human carcinogens. We note that the two tumor types with high CoE in animals in this systematic review are the same as those identified with limited evidence in humans by the IARC Working Group. However, even in cases where the animal evidence demonstrates high CoE, the extrapolation of risk from cancer bioassays to humans is particularly complex for RF EMF. Without a better understanding of the mechanism of the carcinogenicity of RF-EMF, the choice of exposure metric for risk extrapolation (whole body versus localized), intensity or cumulative exposure, whether or not a monotonic dose–response holds for carcinogenic effects, and whether SAR is the appropriate dose metric for adverse effects induced by RF-EMF may be critical.

Other 

This review was partially funded by the WHO radioprotection programme. The protocol for this review was registered in Prospero reg. no. CRD42021265563 and published in Environment International 2022 (Mevissen et al. 2022).

Final conclusions 

The findings of this systematic review indicate that there is evidence that RF EMF exposure increases the incidence of cancer in experimental animals with the CoE being strongest for malignant heart schwannomas and gliomas.

Despite the high level of certainty that evidence of carcinogenicity in experimental animals may predict a carcinogenic hazard to humans, extrapolation of risk from cancer bioassays to humans is particularly complex for RF EMF. Without an understanding of the mechanism of the carcinogenicity of RF-EMF the choice of exposure metric for risk extrapolation (whole body versus localized), intensity or cumulative exposure whether or not a monotonic dose–response holds for carcinogenic effects, and whether SAR is the appropriate dose metric for adverse effects induced by RF-EMF may be critical.

Besides the integration of the sensitivity domain adopted from the RoC, more work is needed to tailor the GRADE approach to assessing the CoE from animal cancer bioassays designed to identifying risks of environmental agents.